Evidence for compensatory upregulation of expressed X-linked genes in mammals, Caenorhabditis elegans and Drosophila melanogaster Public Deposited

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  • Deng, Xinxian
  • Hiatt, Joseph B
  • Nguyen, Di Kim
  • Ercan, Sevinc
  • Sturgill, David
  • Hillier, LaDeana W
  • Schlesinger, Felix
  • Davis, Carrie A
  • Reinke, Valerie J
  • Gingeras, Thomas R
  • Shendure, Jay
  • Waterston, Robert H
  • Oliver, Brian
  • Lieb, Jason D
  • Disteche, Christine M
Abstract
  • Many animal species use a chromosome-based mechanism of sex determination, which has led to the coordinate evolution of dosage-compensation systems. Dosage compensation not only corrects the imbalance in the number of X chromosomes between the sexes but also is hypothesized to correct dosage imbalance within cells that is due to monoallelic X-linked expression and biallelic autosomal expression, by upregulating X-linked genes twofold (termed ‘Ohno’s hypothesis’). Although this hypothesis is well supported by expression analyses of individual X-linked genes and by microarray-based transcriptome analyses, it was challenged by a recent study using RNA sequencing and proteomics. We obtained new, independent RNA-seq data, measured RNA polymerase distribution and reanalyzed published expression data in mammals, C. elegans and Drosophila. Our analyses, which take into account the skewed gene content of the X chromosome, support the hypothesis of upregulation of expressed X-linked genes to balance expression of the genome.
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Resource type
  • Article
Rights statement
  • In Copyright
Journal title
  • Nature Genetics
Journal volume
  • 43
Journal issue
  • 12
Language
  • English
ISSN
  • 1061-4036
  • 1546-1718
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